
Occasional Invader
Harmonia axyridis
The Asian lady beetle, *Harmonia axyridis*, belongs to the family Coccinellidae, order Coleoptera. Adults measure 5.5–8.5 mm in length and exhibit significant color polymorphism, ranging from pale orange to bright red, with 0 to 22 black spots; a characteristic M- or W-shaped black marking on the white pronotum is diagnostic. Its lifecycle comprises four stages: egg, larva (four instars), pupa, and adult, typically completed in 15–30 days depending on temperature and food availability. Females can lay up to 2,000 eggs over their 1–3 month lifespan, with optimal development occurring between 20–25°C and relative humidity of 60–80%. This high reproductive capacity contributes to its rapid establishment and spread in new environments.
Primarily diurnal hunters, *H. axyridis* are voracious predators of soft-bodied arthropods, particularly aphids, psyllids, and scale insects, consuming hundreds daily as both larvae and adults. During late summer and fall, as aphid populations decline and temperatures drop, they exhibit a strong aggregative instinct, releasing aggregation pheromones (e.g., harmoniaxydin) that draw thousands to overwintering sites. When disturbed or stressed, adults reflexively bleed hemolymph – a yellowish, acrid fluid containing toxic alkaloids (harmoniaxydin, harmonine) – from their leg joints. This 'reflex bleeding' stains surfaces and serves as a defense mechanism against predators, while also posing an ophthalmic contact risk to humans in mass aggregations.
As an occasional invader, *H. axyridis* typically seeks harborage in wall voids, attics, soffits, and window/door frames as outdoor temperatures decrease, selecting structures that provide solar exposure and thermal insulation. Their overwintering aggregations are frequently observed on the sunny, south-facing sides of buildings, where they exploit small cracks and crevices for entry; aggregations can number in the thousands or even millions in severe infestations. Foraging patterns during warmer months are primarily arboreal and field-based, following aphid populations. Elevated CO2 levels within structures, along with contrasting light-colored surfaces adjacent to dark entry points, are considered conducive conditions that attract overwintering adults.
Tailored to Asian Lady Beetle